Simulation of multi-species tropical forest dynamics using a vertically and horizontally structured model

Abstract The mutual interaction of forest growth and light conditions causes vertical and horizontal differentiation in the natural forest mosaic. The associated physical processes should be represented in simulation models describing long-term forest dynamics. The FORMIX2 (tropical) forest simulator incorporates the multi-storey canopy layer structure of the natural tropical forest, the basic processes of gap succession dynamics and gap interaction by tree fall and seed dispersal, several functionally different species groups (emergents, main canopy trees, pioneers, woody treelets and forest-floor herbs) and the energy (carbon) balance of assimilation, dissimilation, and increment resulting from the differing light conditions for the various canopy layers in the gap-size areas of a forest mosaic. The vertically and horizontally structured FORMIX2 model produces time-dependent results (average per hectare data) for stem numbers, diameter distributions, basal areas, standing crop and yield, as well as dynamic views of the forest profile and of spatial forest development, as a function of logging method and intensity and silvicultural treatment. The model has been applied in an exploratory investigation to compute the long-term development of (a) virgin forest and (b) logged-over forest in Sabah, using prototypical data. The results obtained agree with empirical data.