New Remains of the Multituberculate Mammal Barbatodon from the Upper Cretaceous of the Hațeg Basin (Romania)

The so-called “Hațeg Island” (Transylvania, Romania) is well known for its rich and peculiar Late Cretaceous vertebrate assemblages, including dwarf dinosaurs, first reported at the end of the 19th century. Besides dinosaurs, other important members of the “Hațeg Island” terrestrial ecosystems were the multituberculate mammals. Late Cretaceous (Maastrichtian) multituberculates are extremely rare in Europe, being known exclusively from the “Hațeg Island.” Two genera are reported from this area: Kogaionon, with the single species K. ungureanui, and Barbatodon, with two species: B. transylvanicus and B. oardaensis. Both genera belong to the European endemic family Kogaionidae. We report herein new remains of B. transylvanicus from the Maastrichtian locality of Pui (Hunedoara District) in the Hațeg Basin, including the most complete dentaries found to date with complete cheek teeth rows. Intraspecific variability of this species in dentary shape, tooth morphology, and size difference is highlighted.