Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane

A new Permian gastropod assemblage from the Roadian (Middle Permian) Khao Khad Formation, Saraburi Group (Lopburi Province, Central Thailand) which is part of the Indochina Terrane, has yielded one of the most diverse Permian gastropod faunas known from Thailand. A total of 44 gastropod species belonging to 30 genera are described herein, including thirteen new species and one new genus. The new genus is Altotomaria. The new species are Bellerophon erawanensis, Biarmeaspira mazaevi, Apachella thailandensis, Gosseletina microstriata, Worthenia humiligrada, Altotomaria reticulata, Yunnania inflata, Trachydomia suwanneeae, Trachyspira eleganta, Heterosubulites longusapertura, Platyzona gradata, Trypanocochlea lopburiensis and Streptacis? khaokhadensis. Most of the species in the studied assemblage represent vetigastropods  (35.6%) and caenogastropods (26.7%) and most of the species belong to Late Palaeozoic cosmopolitan genera. The studied faunas come from shallow water carbonates that are rich in fusulinids, followed by gastropods, ostracods, bivalves and brachiopods. The gastropod assemblage from the Khao Khad Formation shares no species with the gastropod assemblages from other Permian formations in Thailand, the Tak Fa Limestone and the Ratburi Limestone. However, it is similar to the Late Permian gastropod faunas from South China of the Palaeo-Tethys, therefore it suggests that the Indochina Terrane was not located far from South China.