Hepatics from Rovno amber (Ukraine): Frullania pycnoclada and a new species, F. vanae

A study of Rovno amber revealed two species of liverworts in the genus Frullania (Frullaniaceae, Marchantiophyta) that are new to the Late Eocene flora of Rovno amber. Both species are characterized by the mammillose anterior surface of the ventral lobules. One of these, F. pycnoclada Grolle, was described previously from Eocene Baltic amber, whereas the other species, F. vanae Mamontov, J.J.Atwood, Perkovsky & Ignatov, is described here as new. Frullania vanae differs from F. pycnoclada in the shape of the stem and branch underleaves, the shape of the stem leaf lobules, and in the areolation of the leaf lobe cells.