Hippocampus-independent phase precession in entorhinal grid cells

In the brain, both rate and temporal codes are critical for information storage. Theta phase precession is a change in action potential timing in the hippocampus where place cells fire at progressively earlier phases of the theta rhythm as the animal moves across the firing field of the neuron. This paper explores the circuitry of theta phase precession and shows that phase precession is expressed independently of the hippocampus in spatially modulated grid cells in parts of the entorhinal cortex.