Forceps delivery of a new treefrog species of the genus Boophis from eastern Madagascar (Amphibia: Mantellidae)

We describe a new treefrog species of the genus Boophis (subgenus Sahona) from eastern Madagascar. Boophis calcaratus sp. nov. is distinguished from other species of the group by the presence of distinct dermal heel tubercles which are especially well developed in females. We also re-examined the type specimens of Boophis hillenii, a species that has not been rediscovered after its description in 1979. According to our analysis this taxon shows no consistent morphological differences to Boophis idae and supposed bioacoustic differences may be explained by two distinct call types in this species, and we thus suggest to consider hillenii as junior synonym of idae.