Gene Analysis of Sugary and Shrunken Mutants of Rice, Oryza sativa L.

1984 
This study was undertaken to test the allelism of the genes for wrinkled endosperm in rice and to identify the chromosome on which these genes are located. From the results obtained in the F1 and F2, it was concluded that the four genes controlling the wrinkled endosperm character are located at three different loci. The authors propose to designate these four genes as su, shr-1s shr-1a and shr-2, respectively. Segregation ratio in the F2 of the cross between EM-5 (su) and trisomic D was fitted to that of a trisomic segregation. The gene shr-1 was found to be linked to the rolled leaf gene (rl-2) with the recombination value of 23.6%These results suggest that su and shr-1 are located on Chromosome 12 and 3, respectively.
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