A review of aquatic vertebrate remains from the Middle-Upper Triassic Jilh Formation of Saudi Arabia

A recent field survey of the Middle–Upper Triassic (upper Anisian to lowermost Carnian) paralic marine deposits of the Jilh Formation in central Saudi Arabia has yielded large quantities of vertebrate fossils. These finds prompt a revision of the existing faunal list and include at least one novel stratigraphical occurrence for the Arabian Peninsula. The remains comprise sauropterygian marine reptiles (Psephosauriscus sp., Nothosaurus cf. tchernovi, Nothosaurus cf. giganteus, Simosaurus sp.), a lungfish (Ceratodus sp.),hybodontiform sharks (Hybodus sp.) and saurichthyform actinopterygians (Saurichthys sp.). Palaeobiogeographical assessment reinforces Tethyan affinities for the assemblage and reflects the close proximity of the Arabian region to the ‘Sephardic Realm’, a compositionally distinct circum-Mediterranean faunal province characterized by hypersaline Muschelkalk facies.