A five-gene phylogeny of Pezizomycotina

Pezizomycotina is the largest subphylum of Ascomycota and includes the vast majority of filamen- tous, ascoma-producing species. Here we report the results from weighted parsimony, maximum likelihood and Bayesian phylogenetic analyses of five nuclear loci (SSU rDNA, LSU rDNA, RPB1, RPB2 and EF-1a) from 191 taxa. Nine of the 10 Pezizomycotina classes currently recognized were represented in the sam- pling. These data strongly supported the monophyly of Pezizomycotina, Arthoniomycetes, Eurotiomycetes, Orbiliomycetes and Sordariomycetes. Pezizomycetes and Dothideomycetes also were resolved as mono- phyletic but not strongly supported by the data. Lecanoromycetes was resolved as paraphyletic in parsimony analyses but monophyletic in maximum likelihood and Bayesian analyses. Leotiomycetes was polyphyletic due to exclusion of Geoglossaceae. The two most basal classes of Pezizomycotina were Orbilio- mycetes and Pezizomycetes, both of which comprise species that produce apothecial ascomata. The seven