The Nucho-dorsal Glands of Rhabdophis guangdongensis (Squamata: Colubridae: Natricinae), with Notes on Morphological Variation and Phylogeny Based on Additional Specimens

We report the presence of nucho-dorsal glands in Rhabdophis guangdongensis. This recently described species is the 18th member of the genus known to possess integumentary defensive glands. Nine to ten pairs of nuchal glands, separated by a diastema from 116–118 pairs of dorsal glands were observed in two specimens. Because the species was previously known only from the female holotype, we also report data on the size and scalation of five additional specimens and on the morphology of the hemipenis.