Implications of the variety of bat echolocation sounds for understanding biosonar processing

The variety of echolocation sounds used by different species of bats have implications for target ranging. Signals recorded at individual sites reveal species stacked in different frequency bands, perhaps to avoid cross-interference. Search-stage signals include short single-harmonic or multi-harmonic tone-bursts, or very shallow FM bursts. These narrowband sounds have abrupt onsets to evoke phasic on-responses that register echo delay, but with limited acuity. Wider FM sweeps used for searching by other bats evoke on-responses at many more frequencies for better delay acuity. These sound types may signify foraging in the open, within broad spaces bounded relatively remotely by trees or the ground. Intervals between broadcasts are consistent with biosonar operating ranges set by the boundaries of the scene in relation to atmospheric attenuation. Most species make transitions to wider signal bandwidth during interception by increasing FM sweep-width or adding harmonics. Additionally, wideband, multi-harmon...