Floral organogenesis and development of Torenia fournieri.

The flower of Torenia fournieri(Scrophulariaceae)is perfect,usually zygomorphic. Corolla is sympetalous,trumpet shaped and with a lower lip divided into three lobes and a larger upper lip with two lobes. Initiation of five sepal primordia is not consistent in order;The order of initiation is from one adaxial primordium,two abaxial primordia and finally two lateral primordia, while their development is synchronous. Four stamen primordia initiate nearly at the same time before the initiation of the petal primordia and grow slower than the petal primordia. Five petal primordia initiate simultaneously,and two abaxial primordia are fused later. Compared to the floral organogenesis of Agalinis(Scrophulariaceae), there are more differences in the time and order of every floral primordia initiation. While compared to the patterns of floral organogenesis of Rehmannia(Scrophulariaceae),Whytockia,Rhynchoglossum and Titanotrichum(Gesneriaceae),there are not two distinct patterns of the five genera exist in the floral organogenesis,which shows relatively complex relationship between Scrophulariaceae and Gesneriaceae and that the ovary locules and placentation were used to division of these two falimies should be studied further.