Palaeogene and Neogene brachyurans of the Amazon basin: a revised first appearance date for primary freshwater crabs (Brachyura, Trichodactylidae)

We describe claw fragments of fossil primary freshwater crabs from three areas in the Amazon basin, Tarapoto (Early Oligocene) and Contamana (Middle Eocene to early Late Miocene) in Peru, and Eirunepe (Late Miocene) in Brazil. All these fragments most likely belong to the family Trichodactylidae. We show a continuous presence of primary freshwater crabs in proto-Amazonian lowlands from the Middle Eocene to the Late Miocene and can thus shift the earliest appearance date of freshwater-adapted brachyurans into the Eocene, at least in the Neotropics.