Response to Hockley: The merit of economic and biological measures in conservation planning

We welcome Hockley et al.'s response [1] to our recent article in TREE [2] and have few disagreements with its central argument. In particular, we agree that research into the economic benefits provided by natural systems is sorely needed. In our review, we did not intend to advocate that conservation benefits be measured solely in biological terms; we focused on biological measures because they were used by most of the cost studies that we examined. Because our review topic was the cost side of conservation, we are grateful for this opportunity to stress the importance of conservation benefits.