The cattle egret (Bubulcus ibis) is a cosmopolitan species of heron (family Ardeidae) found in the tropics, subtropics, and warm-temperate zones. It is the only member of the monotypic genus Bubulcus, although some authorities regard two of its subspecies as full species, the western cattle egret and the eastern cattle egret. Despite the similarities in plumage to the egrets of the genus Egretta, it is more closely related to the herons of Ardea. Originally native to parts of Asia, Africa, and Europe, it has undergone a rapid expansion in its distribution and successfully colonised much of the rest of the world in the last century. It is a white bird adorned with buff plumes in the breeding season. It nests in colonies, usually near bodies of water and often with other wading birds. The nest is a platform of sticks in trees or shrubs. Cattle egrets exploit drier and open habitats more than other heron species. Their feeding habitats include seasonally inundated grasslands, pastures, farmlands, wetlands, and rice paddies. They often accompany cattle or other large mammals, catching insect and small vertebrate prey disturbed by these animals. Some populations are migratory and others show postbreeding dispersal. The adult cattle egret has few predators, but birds or mammals may raid its nests, and chicks may be lost to starvation, calcium deficiency, or disturbance from other large birds. This species maintains a special relationship with cattle, which extends to other large grazing mammals; wider human farming is believed to be a major cause of their suddenly expanded range. The cattle egret removes ticks and flies from cattle and consumes them. This benefits both species, but it has been implicated in the spread of tick-borne animal diseases. The cattle egret was first described in 1758 by Linnaeus in his Systema naturae as Ardea ibis, but was moved to its current genus by Charles Lucien Bonaparte in 1855. Its genus name Bubulcus is Latin for herdsman, referring, like the English name, to this species' association with cattle. Ibis is a Latin and Greek word which originally referred to another white wading bird, the sacred ibis, but was applied to this species in error. The cattle egret has two geographical races, which are sometimes classified as full species, the western cattle egret, B. ibis, and eastern cattle egret, B. coromandus. The two forms were split by McAllan and Bruce, but were regarded as conspecific by almost all other recent authors until the publication of the influential Birds of South Asia. The eastern subspecies B. (i.) coromandus, described by Pieter Boddaert in 1783, breeds in Asia and Australasia, and the western nominate form occupies the rest of the species range, including the Americas. Some authorities recognise a third Seychelles subspecies, B. i. seychellarum, which was first described by Finn Salomonsen in 1934. Despite superficial similarities in appearance, the cattle egret is more closely related to the genus Ardea, which comprises the great or typical herons and the great egret (A. alba), than to the majority of species termed egrets in the genus Egretta. Rare cases of hybridization with little blue herons Egretta caerulea, little egrets Egretta garzetta and snowy egrets Egretta thula have been recorded. The cattle egret is a stocky heron with an 88–96 cm (35–38 in) wingspan; it is 46–56 cm (18–22 in) long and weighs 270–512 g (9.5–18.1 oz). It has a relatively short, thick neck, a sturdy bill, and a hunched posture. The nonbreeding adult has mainly white plumage, a yellow bill, and greyish-yellow legs. During the breeding season, adults of the nominate western subspecies develop orange-buff plumes on the back, breast, and crown, and the bill, legs, and irises become bright red for a brief period prior to pairing. The sexes are similar, but the male is marginally larger and has slightly longer breeding plumes than the female; juvenile birds lack coloured plumes and have a black bill. B. i. coromandus differs from the nominate subspecies in breeding plumage, when the buff colour on its head extends to the cheeks and throat, and the plumes are more golden in colour. This subspecies' bill and tarsus are longer on average than in B. i. ibis. B. i. seychellarum is smaller and shorter-winged than the other forms. It has white cheeks and throat, like B. i. ibis, but the nuptial plumes are golden, as with B. i. coromandus. Individuals with abnormally grey, melanistic plumages have been recorded.