The atrium (Latin ātrium, “entry hall”), is the upper chamber through which blood enters the ventricles of the heart. There are two atria in the human heart – the left atrium receives blood from the pulmonary (lung) circulation, and the right atrium receives blood from the venae cavae (venous circulation). The atria receive blood while relaxed (diastole), then contract (systole) to move blood to the ventricles. All animals with a closed circulatory system have at least one atrium. Humans have two atria. The atrium (Latin ātrium, “entry hall”), is the upper chamber through which blood enters the ventricles of the heart. There are two atria in the human heart – the left atrium receives blood from the pulmonary (lung) circulation, and the right atrium receives blood from the venae cavae (venous circulation). The atria receive blood while relaxed (diastole), then contract (systole) to move blood to the ventricles. All animals with a closed circulatory system have at least one atrium. Humans have two atria. The atrium was formerly called the 'auricle'; that term is still used to describe this chamber in some other animals, such as the Mollusca. Humans have a four-chambered heart consisting of the right atrium, left atrium, right ventricle, and left ventricle. The atria are the two upper chambers. The right atrium receives and holds deoxygenated blood from the superior vena cava, inferior vena cava, anterior cardiac veins and smallest cardiac veins and the coronary sinus, which it then sends down to the right ventricle (through the tricuspid valve), which in turn sends it to the pulmonary artery for pulmonary circulation. The left atrium receives the oxygenated blood from the left and right pulmonary veins, which it pumps to the left ventricle (through the mitral valve) for pumping out through the aorta for systemic circulation. The right atrium and right ventricle are often referred to as the right heart; similarly, the left atrium and left ventricle are often referred to as the left heart. The atria do not have valves at their inlets, and as a result, a venous pulsation is normal and can be detected in the jugular vein as the jugular venous pressure. Internally, there are the rough pectinate muscles and crista terminalis of His, which act as a boundary inside the atrium and the smooth-walled part of the right atrium, the sinus venarum, which are derived from the sinus venosus. The sinus venarum is the adult remnant of the sinus venous and it surrounds the openings of the venae cavae and the coronary sinus. Attached to the right atrium is the right atrial appendage – a pouch-like extension of the pectinate muscles. The interatrial septum separates the right atrium from the left atrium; this is marked by a depression in the right atrium – the fossa ovalis. The atria are depolarised by calcium. High in the upper part of the left atrium is a muscular ear-shaped pouch – the left atrial appendage. This appears to 'function as a decompression chamber during left ventricular systole and during other periods when left atrial pressure is high'. The sinoatrial (SA) node is located in the posterior aspect of the right atrium, next to the superior vena cava. This is a group of pacemaker cells which spontaneously depolarize to create an action potential. The cardiac action potential then spreads across both atria causing them to contract, forcing the blood they hold into their corresponding ventricles. The atrioventricular node (AV node) is another node in the cardiac electrical conduction system. This is located between the atria and the ventricles. The left atrium is supplied mainly by the left circumflex coronary artery, and its small branches. The oblique vein of the left atrium is partly responsible for venous drainage; it derives from the embryonic left superior vena cava.