Beech marten

The beech marten (Martes foina), also known as the stone marten, house marten or white breasted marten, is a species of marten native to much of Europe and Central Asia, though it has established a feral population in North America. It is listed as Least Concern by the IUCN on account of its wide distribution, its large population, and its presence in a number of protected areas. It is superficially similar to the pine marten, but differs from it by its smaller size and habitat preferences. While the pine marten is a forest specialist, the beech marten is a more generalist and adaptable species, occurring in a number of open and forest habitats. Its most likely ancestor is Martes vetus, which also gave rise to the pine marten. The earliest M. vetus fossils were found in deposits dated to the Würm glaciation in Lebanon and Israel. The beech marten likely originated in the Near East or southwestern Asia, and may have arrived in Europe by the Late Pleistocene or the early Holocene. Thus, the beech marten differs from most other European mustelids of the Quaternary, as all other species (save for the European mink) appeared during the Middle Pleistocene. Comparisons between fossil animals and their descendants indicate that the beech marten underwent a decrease in size beginning in the Würm period. Beech martens indigenous to the Aegean Islands represent a relic population with primitive Asiatic affinities. The skull of the beech marten suggests a higher adaptation than the pine marten toward hypercarnivory, as indicated by its smaller head, shorter snout and its narrower post-orbital constriction and lesser emphasis on cheek teeth. Selective pressures must have acted to increase the beech marten's bite force at the expense of gape. These traits probably acted on male beech martens as a mechanism to avoid both intraspecific competition with females and interspecific competition with the ecologically overlapping pine marten. As of 2005, eleven subspecies are recognised. The beech marten is superficially similar to the pine marten, but has a somewhat longer tail, a more elongated and angular head and has shorter, more rounded and widely spaced ears. Its nose is also of a light peach or grey colour, whereas that of the pine marten is dark black or greyish-black. Its feet are not as densely furred as those of the pine marten, thus making them look less broad, with the paw pads remaining visible even in winter. Because of its shorter limbs, the beech marten's manner of locomotion differs from that of the pine marten; the beech marten moves by creeping in a polecat-like manner, whereas the pine marten and sable move by bounds. The weight load per 1 cm2 of the supporting surface of the beech marten's foot (30.9 gm) is double that of the pine marten (15.2 gm), thus it is obliged to avoid snowy regions. Its skull is similar to that of the pine marten, but differs in its shorter facial region, more convex profile, its larger carnassials and smaller molars. The beech marten's penis is larger than the pine marten's, with the bacula of young beech martens often outsizing those of old pine martens. Males measure 430–590 mm in body length, while females measure 380–470 mm. The tail measures 250–320 mm in males and 230–275 mm in females. Males weigh 1.7–1.8 kg in winter and 2–2.1 kg in summer, while females weigh 1.1–1.3 kg in winter and 1.4–1.5 kg in summer. The beech marten's fur is coarser than the pine marten's, with elastic guard hairs and less dense underfur. Its summer coat is short, sparse and coarse, and the tail is sparsely furred. The colour tone is lighter than the pine marten's. Unlike the pine marten, its underfur is whitish, rather than greyish. The tail is dark-brown, while the back is darker than that of the pine marten. The throat patch of the beech marten is always white. The patch is large and generally has two projections extending backwards to the base of the forelegs and upward on the legs. The dark colour of the belly juts out between the forelegs as a line into the white colour of the chest and sometimes into the neck. In the pine marten, by contrast, the white colour between the forelegs juts backwards as a protrusion into the belly colour. The beech marten is mainly a crepuscular and nocturnal animal, though to a much lesser extent than the European polecat. It is especially active during moonlit nights. Being a more terrestrial animal than the pine marten, the beech marten is less arboreal in its habits, though it can be a skilled climber in heavily forested areas. It is a skilled swimmer, and may occasionally be active during daytime hours, particularly in the summer, when nights are short. It typically hunts on the ground. During heavy snowfalls, the beech marten moves through paths made by hares or skis.