Antbird

The antbirds are a large passerine bird family, Thamnophilidae, found across subtropical and tropical Central and South America, from Mexico to Argentina. There are more than 230 species, known variously as antshrikes, antwrens, antvireos, fire-eyes, bare-eyes and bushbirds. They are related to the antthrushes and antpittas (family Formicariidae), the tapaculos, the gnateaters and the ovenbirds. Despite some species' common names, this family is not closely related to the wrens, vireos or shrikes. Antbirds are generally small birds with rounded wings and strong legs. They have mostly sombre grey, white, brown and rufous plumage, which is sexually dimorphic in pattern and colouring. Some species communicate warnings to rivals by exposing white feather patches on their backs or shoulders. Most have heavy bills, which in many species are hooked at the tip. Most species live in forests, although a few are found in other habitats. Insects and other arthropods form the most important part of their diet, although small vertebrates are occasionally taken. Most species feed in the understory and midstory of the forest, although a few feed in the canopy and a few on the ground. Many join mixed-species feeding flocks, and a few species are core members. To various degrees, around eighteen species specialise in following swarms of army ants to eat the small invertebrates flushed by the ants, and many others may feed in this way opportunistically. Antbirds are monogamous, mate for life, and defend territories. They usually lay two eggs in a nest that is either suspended from branches or supported on a branch, stump, or mound on the ground. Both parents share the tasks of incubation and of brooding and feeding the nestlings. After fledging, each parent cares exclusively for one chick. Thirty-eight species are threatened with extinction as a result of human activities. Antbirds are not targeted by either hunters or the pet trade. The principal threat is habitat loss, which causes habitat fragmentation and increased nest predation in habitat fragments. The antbird family Thamnophilidae used to be considered a subfamily, Thamnophilinae, within a larger family Formicariidae that included antthrushes and antpittas. Formerly, that larger family was known as the 'antbird family' and the Thamnophilinae were 'typical antbirds'. In this article, 'antbird' and 'antbird family' refer to the family Thamnophilidae. Thamnophilidae was removed from Formicariidae, leaving behind the antthrushes and antpittas, due to recognition of differences in the structure of the breastbone (sternum) and syrinx, and Sibley and Ahlquist's examination of DNA–DNA hybridization. The Thamnophilidae antbirds are members of the infraorder Tyrannides (or tracheophone suboscines), one of two infraorders in the suborder Tyranni. The Thamnophilidae are now thought to occupy a fairly basal position within the infraorder, i. e. with regard to their relatives the antthrushes and antpittas, tapaculos, gnateaters, and also the ovenbirds. The sister group of the Thamnophilidae is thought to be the gnateaters. The ovenbirds, tapaculos, antthrushes and antpittas are thought to represent a different radiation of that early split. The antbird family contains over 230 species, variously called antwrens, antvireos, antbirds and antshrikes. The names refer to the relative sizes of the birds (increasing in the order given, though with exceptions) rather than any particular resemblance to the true wrens, vireos or shrikes. In addition, members of the genus Phlegopsis are known as bare-eyes, Pyriglena as fire-eyes and Neoctantes and Clytoctantes as bushbirds. Although the systematics of the Thamnophilidae is based on studies from the mid-19th century, when fewer than half the present species were known, comparison of the myoglobin intron 2, GAPDH intron 11 and the mtDNA cytochrome b DNA sequences has largely confirmed it. There are two major clades – most antshrikes and other larger, strong-billed species as well as Herpsilochmus, versus the classical antwrens and other more slender, longer-billed species – and the monophyly of most genera was confirmed.