Amelanchier

Amelanchier (/æməˈlænʃɪər/ am-ə-LAN-sheer), also known as shadbush, shadwood or shadblow, serviceberry or sarvisberry, or just sarvis, juneberry, saskatoon, sugarplum or wild-plum, and chuckley pear is a genus of about 20 species of deciduous-leaved shrubs and small trees in the rose family (Rosaceae). Amelanchier is native to temperate regions of the Northern Hemisphere, growing primarily in early successional habitats. It is most diverse taxonomically in North America, especially in the northeastern United States and adjacent southeastern Canada, and at least one species is native to every U.S. state except Hawaii and to every Canadian province and territory. Two species also occur in Asia, and one in Europe. The taxonomic classification of shadbushes has long perplexed botanists, horticulturalists, and others, as suggested by the range in number of species recognized in the genus, from 6 to 33, in two recent publications. A major source of complexity comes from the occurrence of hybridization, polyploidy, and apomixis (asexual seed production), making species difficult to characterize and identify. The various species of Amelanchier grow to 0.2–20 m tall; some are small trees, some are multistemmed, clump-forming shrubs, and yet others form extensive low shrubby patches (clones). The bark is gray or less often brown, and in tree species smooth or fissuring when older. The leaves are deciduous, cauline, alternate, simple, lanceolate to elliptic to orbiculate, 0.5–10 x 0.5–5.5 cm, thin to coriaceous, with surfaces above glabrous or densely tomentose at flowering, and glabrous or more or less hairy beneath at maturity. The inflorescences are terminal, with 1–20 flowers, erect or drooping, either in clusters of one to four flowers, or in racemes with 4–20 flowers. The flowers have five white (rarely somewhat pink, yellow, or streaked with red), linear to orbiculate petals, 2.6–25 mm long, with the petals in one species (A. nantucketensis) often andropetalous (bearing apical microsporangia adaxially). The flowers appear in early spring, 'when the shad run' according to North-American tradition (leading to names such as 'shadbush'). The fruit is a berry-like pome, red to purple to nearly black at maturity, 5–15 mm diameter, insipid to delectably sweet, maturing in summer. Amelanchier plants are valued horticulturally, and their fruits are important to wildlife. For North American species, the taxonomy follows the Flora of North America; for Asian species the Flora of China; and for the one European species the Flora Europaea. Since classifications have varied greatly over the past century, species names are often used interchangeably in the nursery trade. Several natural or horticultural hybrids also exist, and many A. arborea and A. canadensis plants that are offered for sale are actually hybrids, or entirely different species. A. × grandiflora is another hybrid of garden origin, between A. arborea and A. laevis. The cultivar 'La Paloma' has gained the Royal Horticultural Society’s Award of Garden Merit. A taxon called Amelanchier lamarckii (or A. x lamarckii) is very widely cultivated and naturalized in Europe, where it was introduced in the 17th century. It is apomictic, breeding true from seed, and probably of hybrid origin, perhaps descending from a cross between A. laevis and either A. arborea or A. canadensis. While A. lamarckii is known to be of North American origin, probably from eastern Canada, it is not known to occur naturally in the wild in North America. The origin of the generic name Amelanchier is probably derived from amalenquièr, amelanchièr, the Provençal names of the European Amelanchier ovalis.