Vagaceratops

Vagaceratops (meaning 'wandering (vagus, Latin) horned face', in reference to its close relationship with Kosmoceratops from Utah) is a genus of herbivorous ceratopsian dinosaur. It is a chasmosaurine ceratopsian which lived during the Late Cretaceous period (late Campanian) in what is now Alberta. Its fossils have been recovered from the Upper Dinosaur Park Formation. Vagaceratops is known primarily from three fossil skulls. Although the general structure was typical of ceratopsids (i.e. a parrot-like beak, large neck frill, and nasal horn) it has some peculiarities. The skulls are characterized by a reduced supraorbital horn, brow horns that are reduced to low bosses and a larger snout compared to related animals. Vagaceratops had smaller parietal fenestrae than most ceratopsids and had a strange configuration of epoccipitals (bones surrounding the frill). It possessed ten epoccipitals, eight of which were centrally flattened, curved forward and upward and fused together to form a jagged margin along the back of the frill. The frill was shorter and more square-shaped than other chasmosaurines, being wider than it was long. Vagaceratops was named by Scott D. Sampson, Mark A. Loewen, Andrew A. Farke, Eric M. Roberts, Catherine A. Forster, Joshua A. Smith, and Alan L. Titus in 2010, and the type species is Vagaceratops irvinensis. This species was originally described as a species of Chasmosaurus (C. irvinensis) in 2001. Its relationships remain debated. Vagaceratops has variously been allied with Kosmoceratops or with Chasmosaurus. The cladogram below is the phylogeny of the Chasmosaurinae by Brown et al (2015):