Arctica islandica

The ocean quahog (Arctica islandica) is a species of edible clam, a marine bivalve mollusk in the family Arcticidae. This species is native to the North Atlantic Ocean, and it is harvested commercially as a food source. This species is also known by a number of different common names, including Icelandic cyprine, mahogany clam, mahogany quahog, black quahog, and black clam. The typical Arctica islandica resembles the quahog, but the shell of the ocean quahog is rounder, the periostracum is usually black, and on the interior of the shell, the pallial line has no indentation, or sinus. Unlike the quahog, which lives intertidally and can be collected by clam digging, this species lives subtidally, and can only be collected by dredging. They grow to sizes exceeding 50 mm shell height. An individual specimen was reported to have lived 507 years, making it the longest-lived non-colonial metazoan whose age was accurately known. These animals show exceptional longevity with a highest reported age, for Ming the clam, of 507 years. This makes it the longest living non-colonial metazoan species with an authenticated lifespan. It is unknown how long it could have lived if it had not been collected alive by an expedition in 2006. In 1868 one specimen, collected alive near Iceland, was 374 years old. The study of its growth rate and the oxygen isotope data showed that it had a highly variable growth at the peak of the Little Ice Age around 1550–1620 and mild climate near its end around 1765–1780 and had recorded the volcanic eruption of Mount Tambora in 1815. One study found that in animals aged 4–192 years, antioxidant enzymes declined rapidly in the first 25 years, which includes the growth and sexual maturity stages, but afterwards remained stable for over 150 years. Though more detailed studies are warranted, it appears this species is a case of negligible senescence. In contrast to the exceptionally long-lived populations in relatively deep, cold parts of its range, more southern populations that experience greater seasonal variations in salinity and temperature are typically far shorter-lived. For example, A. islandica from the German Bay of Kiel typically only reach an age of about 30 years and those from the German Bight about 150. According to the research done by Irene Ballesta-Artero, Starrlight Augustine, et al,. they conducted experiments using a Dynamic Energy Model (DEM) to test the extreme longevity of the A. Islandica. The A. Islandica species shows slower growth rate than other species of clams, it takes an average rate of 4-days for birth (embryo to larvae stage). Furthermore it takes an average of 5.8-years for the Arctica Islandica to reach maturity, at this point the somatic costs start going down. A. Islandica's extreme longevity arises from a lowered somatic maintenance costs and a low aging acceleration.