The red flour beetle (Tribolium castaneum) is a species of beetle in the family Tenebrionidae, the darkling beetles. It is a worldwide pest of stored products, particularly food grains, and a model organism for ethological and food safety research. The red flour beetle attacks stored grain and other food products including flour, cereals, pasta, biscuits, beans, and nuts, causing loss and damage. The United Nations, in a recent post-harvest compendium, estimated that Tribolium castaneum and Tribolium confusum, the confused flour beetle, are 'the two most common secondary pests of all plant commodities in store throughout the world.' The red flour beetle is of Indo-Australian origin and less able to survive outdoors than the closely related species Tribolium confusum. It has, as a consequence, a more southern distribution, though both species are worldwide in heated environments. The adult is long-lived, sometimes living more than three years. Although previously regarded as a relatively sedentary insect, it has been shown in molecular and ecological research to disperse considerable distances by flight.