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Fish locomotion

Fish locomotion is the variety of types of animal locomotion used by fish, principally by swimming. This however is achieved in different groups of fish by a variety of mechanisms of propulsion in water, most often by wavelike movements of the fish's body and tail, and in various specialised fish by movements of the fins. The major forms of locomotion in fish are anguilliform, in which a wave passes evenly along a long slender body; sub-carangiform, in which the wave increases quickly in amplitude towards the tail; carangiform, in which the wave is concentrated near the tail, which oscillates rapidly; thunniform, rapid swimming with a large powerful crescent-shaped tail; and ostraciiform, with almost no oscillation except of the tail fin. More specialised fish include movement by pectoral fins with a mainly stiff body, as in the sunfish; and movement by propagating a wave along the long fins with a motionless body in fish with electric organs such as the knifefish. In addition, some fish can variously 'walk', i.e., move over land, burrow in mud, and glide through the air. Fish swim by exerting force against the surrounding water. There are exceptions, but this is normally achieved by the fish contracting muscles on either side of its body in order to generate waves of flexion that travel the length of the body from nose to tail, generally getting larger as they go along. The vector forces exerted on the water by such motion cancel out laterally, but generate a net force backwards which in turn pushes the fish forward through the water. Most fishes generate thrust using lateral movements of their body and caudal fin, but many other species move mainly using their median and paired fins. The latter group swim slowly, but can turn rapidly, as is needed when living in coral reefs for example. But they can't swim as fast as fish using their bodies and caudal fins. There are five groups that differ in the fraction of their body that is displaced laterally: In the anguilliform group, containing some long, slender fish such as eels, there is little increase in the amplitude of the flexion wave as it passes along the body. The sub-carangiform group has a more marked increase in wave amplitude along the body with the vast majority of the work being done by the rear half of the fish. In general, the fish body is stiffer, making for higher speed but reduced maneuverability. Trout use sub-carangiform locomotion. The carangiform group, named for the Carangidae, are stiffer and faster-moving than the previous groups. The vast majority of movement is concentrated in the very rear of the body and tail. Carangiform swimmers generally have rapidly oscillating tails. The thunniform group contains high-speed long-distance swimmers, and is characteristic of tunas and is also found in several lamnid sharks. Here, virtually all the sideways movement is in the tail and the region connecting the main body to the tail (the peduncle). The tail itself tends to be large and crescent shaped. This form of swimming enables these fish to chase and catch prey easier due to the increase in speed of swimming, like in Barracudas.

[ "Kinematics", "Fin", "Fish <Actinopterygii>", "Chlopsidae", "Nettastomatidae" ]
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