Western gray squirrel

The western gray squirrel (Sciurus griseus) is an arboreal rodent found along the western coast of the United States and Mexico. It is a tree squirrel. In some places, this species has also been known as the silver-gray squirrel, the California gray squirrel, the Oregon gray squirrel, the Columbian gray squirrel and the banner-tail. There are three geographical subspecies: Sciurus griseus griseus (central Washington to the western Sierra Nevada in central California); S. g. nigripes (from south of San Francisco Bay to San Luis Obispo County, California); and S. g. anthonyi (which ranges from San Luis Obispo to northern Baja California). The western gray squirrel was first described by George Ord in 1818 based on notes taken by Lewis and Clark at The Dalles in Wasco County, Oregon. Sciurus griseus is the largest tree squirrel in the Sierra Nevada and Central California range. It has plantigrade, pentadactyl feet with two phalanges. Compared with the eastern gray squirrel S. carolinensis or the fox squirrel S. niger (which have been introduced into its native range), these squirrels are shy, and will generally run up a tree and give a hoarse chirping call when disturbed. Weights vary from about .35 to 1 kilogram (0.77 to 2.20 lb), and length (including tail) from 43 to 61 centimetres (17 to 24 in). It is the largest native tree squirrel in the western coastal United States. Western gray squirrels exhibit a form of coloration known as counter shading. The dorsal fur is a silver gunmetal gray, with pure white on the underside; there may be black flecks in the tail. Ears are large but without tufts. The ears turn reddish-brown at the back in the winter. The tail is long and typically very bushy. Also, it stays in a curved upwards in an 'S' shape. Tree squirrels undergo a complete head-to-tail molt in the spring and a rump-to-head molt in the fall. Tail hair is replaced only in the spring. Nesting mothers will use their tail hair to line birthing nests. Western gray squirrels eat berries, nuts, a variety of seeds, and the eggs of small birds. The dental formula for Sciurus griseus is 1.0.2.31.0.1.3 × 2 = 22. Western gray squirrels reach sexual maturity at 10 to 11 months, and at approximately one year of age, will begin breeding. They mate over an extended period ranging from December through June. Young are born after approximate 44-day gestation period. Juveniles emerge from nests between March and mid-August. Litter sizes range from one to five kits which remain in the nest for a longer period than other squirrels. The kits are relatively slow in development, and will not leave the nest for six months or more, another species disadvantage when in competition with other, more-rapidly fledging squirrels. Young gray squirrels have furled tails which will not reach fullness until adulthood. This is a good indicator of age and maturity. Mother squirrels often seem to be overworked with a stressed appearance, complete with bruised and battered nipples. Mating squirrels can be very physical and will bite and injure each other. Females can be quite territorial, and will chase others away and have fairly violent altercations between themselves. Western gray squirrels are forest dwellers, and can be found at elevations up to 2,000 m. Time on the ground is spent foraging, but they prefer to travel distances from tree to tree. They are strictly diurnal, and feed mainly on seeds and nuts, particularly pine seeds and acorns, though they will also take berries, fungus and insects. Pine nuts and acorns are considered critical foods because they are very high in oil and moderately high in carbohydrates, which help increase the development of body fat. They feed mostly in trees and on the ground. They generally forage in the morning and late afternoon for acorns, pine nuts, new tree buds, and fruits. They feed on pinecones and many other nuts in preparation for the winter. When on alert, they will spread their tails lavishly, creating an umbrella effect that shields them and possibly provides cover from overhead predators. They are scatter-hoarders making numerous caches of food when it is abundant, and thus contribute to the seed dispersion of their food trees. Although squirrels show relatively good scent relocation abilities, some food caches are never reclaimed, becoming seedlings in the spring. Though they do not hibernate, they do become less active during the winter. Like many prey animals, they depend on auditory alerts from other squirrels or birds to determine safety. Once an alarm call is transmitted, those present will join in, and the trees become a cacophony of chirping squirrels. Tree squirrels are prey for bobcats, hawks, eagles, mountain lions, coyotes, cats, raccoons and humans.