Pinus longaeva

Pinus longaeva (commonly referred to as the Great Basin bristlecone pine, intermountain bristlecone pine, or western bristlecone pine) is a long-living species of bristlecone pine tree found in the higher mountains of California, Nevada, and Utah. Methuselah is a bristlecone pine that is 4,851 years old and the oldest known living non-clonal organism on Earth. In 1987, the bristlecone pine was designated one of Nevada's state trees. It is a medium-size tree, reaching 5 to 15 m (16 to 49 ft) tall and with a trunk diameter of up to 2.5 to 3.6 m (8 to 12 ft). The bark is bright orange-yellow, thin and scaly at the base of the trunk. The needles are in fascicles of five, stout, 2.5 to 4 cm (1 to 1 1⁄2 in) long, deep green to blue-green on the outer face, with stomata confined to a bright white band on the inner surfaces. The leaves show the longest persistence of any plant, with some remaining green for 45 years (Ewers & Schmid 1981). The cones are ovoid-cylindrical, 5 to 10 cm (2 to 4 in) long and 3 to 4 cm (1 to 1 1⁄2 in) broad when closed, green or purple at first, ripening orange-buff when 16 months old, with numerous thin, fragile scales, each scale with a bristle-like spine 2 to 5 mm (1⁄16 to 3⁄16 in) long. The cones open to 4 to 6 cm (1 1⁄2 to 2 1⁄2 in) broad when mature, releasing the seeds immediately after opening. The seeds are 5 mm (3⁄16 in) long, with a 12 to 22 mm (1⁄2 to 7⁄8 in) wing; they are mostly dispersed by the wind, but some are also dispersed by Clark's nutcrackers. These ancient trees have a gnarled and stunted appearance, especially those found at high altitudes, and have reddish-brown bark with deep fissures. As the tree ages, much of its vascular cambium layer may die. In very old specimens, often only a narrow strip of living tissue connects the roots to a handful of live branches. The Great Basin bristlecone pine differs from the Rocky Mountain bristlecone pine in that the needles of the former always have two uninterrupted resin canals, so it lacks the characteristic small white resin flecks appearing on the needles of the latter. The Great Basin bristlecone pine differs from the foxtail pine because the cone bristles of the former are over 2 mm (0.079 in) long, and the cones have a more rounded (not conic) base. The green pine needles give the twisted branches a bottle-brush appearance. The name 'bristlecone pine' refers to the dark purple female cones that bear incurved prickles on their surface. The species occurs in Utah, Nevada and eastern California. In California, it is restricted to the White Mountains, the Inyo Mountains, and the Panamint Range, in Mono and Inyo counties. In Nevada, it is found in most of the higher ranges of the Basin and Range from the Spring Mountains near Las Vegas north to the Ruby Mountains, and in Utah, northeast to South Tent in the Wasatch Range. Due to the inaccessibility of many of the sites that this species occurs, information on their location and abundance is incomplete, and thus is needed.  Environmental niche modelling has been used to better map the distribution of Great Basin bristlecone pine using topographic and spectral variables calculated from a geographic information system (GIS). The tree grows in large open stands, unlike the related foxtail pine, which sometimes form dense forests. Pinus longaeva trees generally do not form closed canopies, usually covering only 15-50%. Pinus longaeva shares habitats with a number of other pine species, including the ponderosa pine, the white fir and, notably, the limber pine, a similarly long-lived high-elevation species. The tree is a 'vigorous' primary succession species, growing quickly on new open ground. It is a 'poor competitor' in good soils, however, and the species does best in harsh terrain. Pinus longaeva is often the dominant species in high-elevation dolomite soils, where few plants can grow. Bristlecone pines are protected in a number of areas owned by the United States federal government, such as the Ancient Bristlecone Pine Forest in the White Mountains of California and the Great Basin National Park in Nevada. These areas prohibit the cutting or gathering of wood.