The white-bellied sea eagle (Haliaeetus leucogaster), also known as the white-breasted sea eagle, is a large diurnal bird of prey in the family Accipitridae. Originally described by Johann Friedrich Gmelin in 1788, it is closely related to Sanford's sea eagle of the Solomon Islands, and the two are considered a superspecies. A distinctive bird, the adult white-bellied sea eagle has a white head, breast, under-wing coverts and tail. The upper parts are grey and the black under-wing flight feathers contrast with the white coverts. The tail is short and wedge-shaped as in all Haliaeetus species. Like many raptors, the female is slightly larger than the male, and can measure up to 90 cm (35 in) long with a wingspan of up to 2.2 m (7.2 ft), and weigh 4.5 kg (9.9 lb). Immature birds have brown plumage, which is gradually replaced by white until the age of five or six years. The call is a loud goose-like honking. Resident from India and Sri Lanka through Southeast Asia to Australia on coasts and major waterways, the white-bellied sea eagle breeds and hunts near water, and fish form around half of its diet. Opportunistic, it consumes carrion and a wide variety of animals. Although rated as Least Concern globally, it has declined in parts of southeast Asia such as Thailand, and southeastern Australia. It is ranked as Threatened in Victoria and Vulnerable in South Australia and Tasmania. Human disturbance to its habitat is the main threat, both from direct human activity near nests which impacts on breeding success, and from removal of suitable trees for nesting. The white-bellied sea eagle is revered by indigenous people in many parts of Australia, and is the subject of various folk tales throughout its range. The white-bellied sea eagle was first described by the German naturalist Johann Friedrich Gmelin in 1788, although John Latham had made notes on the species in 1781, from a specimen obtained in February 1780 at Princes Island off the westernmost cape of Java during Captain Cook's last voyage. Its specific name is derived from the Ancient Greek leuko- 'white', and gaster 'belly'. Its closest relative is the little-known Sanford's sea eagle of the Solomon Islands. These form a superspecies, and as is usual in other sea eagle superspecies, one (the white-bellied sea eagle) has a white head, as opposed to the other species' dark head. The bill and eyes are dark, and the talons are dark yellow as in all Southern Hemisphere sea eagles. Both these species have at least some dark colouration in their tails, though this may not always be clearly visible in the white-bellied sea eagle. The nucleotide sequences of the cytochrome b gene of the two sea eagles were among those analysed in a 1996 study. Although they differ greatly in appearance and ecology, their genetic divergence of 0.3% indicates that the ancestors of the two forms might have diverged as recently as 150,000 years ago. The study authors conclude that although the genetic divergence is more consistent with subspecies, the distinctness in appearance and behaviour warrants the two being retained as separate species. Mitochondrial sequence of the cytochrome b locus differs very slightly from that of Sanford's sea eagle suggesting a relatively recent divergence after New Guinea-based white-bellied sea eagles colonised the Solomon Islands. The white-bellied sea eagle's affinities beyond the Sanford's sea eagle are a little less clear; molecular data indicate that it is one of four species of tropical sea eagle (along with the African fish eagle and the Madagascar fish eagle), while allozyme data indicate it might have a closer relationship with the sea eagles of the northern hemisphere. A further molecular study published in 2005 showed the white-bellied and Sanford's sea eagles to be basal to the four fish eagles (the two mentioned above plus the two hitherto untested species of the genus Ichthyophaga). As well as white-bellied sea eagle and white-breasted sea eagle, other recorded names include white-bellied fish-hawk, white-eagle, and grey-backed sea eagle. The white-bellied sea eagle has a white head, rump and underparts, and dark or slate-grey back and wings. In flight, the black flight feathers on the wings are easily seen when the bird is viewed from below. The large, hooked bill is a leaden blue-grey with a darker tip, and the irides are dark brown. The cere is also lead grey. The legs and feet are yellow or grey, with long black talons (claws). Unlike those of eagles of the genus Aquila, the legs are not feathered. The sexes are similar. Males are 66–80 cm (26–31 in) long and weigh 1.8–3 kg (4.0–6.6 lb). Females are slightly larger, at 80–90 cm (31–35 in) and 2.5–4.5 kg (5.5–9.9 lb). The wingspan ranges from 1.78 to 2.2 m (5.8 to 7.2 ft). A 2004 study on 37 birds from Australia and Papua New Guinea (3 °S to 50 °S) found that birds could be sexed reliably on size, and that birds from latitudes further south were larger than those from the north. There is no seasonal variation in plumage. The moulting pattern of the white-bellied sea eagle is poorly known. It appears to take longer than a year to complete, and can be interrupted and later resumed from the point of interruption. The wings are modified when gliding so that they rise from the body at an angle, but are closer to horizontal further along the wingspan. In silhouette, the comparatively long neck, head and beak stick out from the front almost as far as the tail does behind. For active flight, the white-bellied sea eagle alternates strong deep wing-beats with short periods of gliding. A young white-bellied sea eagle in its first year is predominantly brown, with pale cream-streaked plumage on their head, neck, nape and rump areas. The plumage becomes more infiltrated with white until it acquires the complete adult plumage by the fourth or fifth year. The species breeds from around six years of age onwards. The lifespan is thought to be around 30 years.