Pecten maximus

Pecten maximus, common names the great scallop, king scallop, St James shell or escallop, is a northeast Atlantic species of scallop, an edible saltwater clam, a marine bivalve mollusc in the family Pectinidae. This is the type species of the genus. This species may be conspecific with Pecten jacobaeus, the pilgrim's scallop, which has a much more restricted distribution. The shell of Pecten maximus is quite robust and is characterised by having 'ears' of equal size on either side of the apex. The right, or lower, valve is convex and slightly overlaps the flat left, or upper, valve, which is flat. Larger specimens have an nearly circular outline and the largest may measure 21 cm in length. The 'ears' are prominent and are a minimum of half the width of the shell with the byssal notch situated in the right anterior ear being slight and not serrated. The sculpture of the valves is distinctive and consists of 12 to 17 wide radiating ribs and numerous concentric lines which clearly show the scallop's growth history, while the 'ears' show a few thin ribs which radiate from the beaks. The radiating ribs reach the margins of the valves and this creates a crenulated form. The left valve is normally reddish-brown while the right valve varies from white through cream to shades of pale brown contrasting with pink, red or pale yellow tints; either valve may show zigzag patterns and may also show bands and spots of red, pink or bright yellow. The colour of the body of Pecten maximus is pink or red with the mantle marbled brown and white. When young they are attached to the substrate by a byssus but mature animals are capable of swimming by the opening and rapid closing of the valves. The adductor muscle which is used to close and open the valves is very large and powerful. The foot is a finger-like organ, which spins the byssal threads, which pass through the byssal notch on the ears. The margin of the mantle has two layers the inner layer is finely fringed while the outer is lined with long tentacles with two series totalling 30-36 dark blue or green simple eyes or ocelli in two rows at their base. Pecten maximus occurs in the eastern Atlantic along the European coast from northern Norway, south to the Iberian peninsula, it has also been reported off West Africa, off the Macaronesian Islands. In Great Britain and Ireland it is distributed all round the coast but it is uncommon and localised on the eastern North Sea coast. It prefers offshore waters down to 100m Pecten maximus frequently creates a slight hollow in the substrate for its shell to lie in by opening and closing the valve to eject water from the mantle cavity, which raises the shell at an angle to the substrate so that subsequent water jets into the sediment and create a recess. Once settled, sand, mud, gravel or living organisms coat the upper valve, and the margin of the shell, with the tentacles and eyes, is all that is visible. They are filter feeders which extract particles from the surrounding water via a feeding current which is drawn by cilia across the gills where the food particles are trapped, then taken to the mouth in a stream of mucous. Pecten maximus swims but this is generally limited to escape reactions. The main predators which cause this reaction when detected are the mollusc eating starfish Asterias rubens and Astropecten irregularis, although starfish which do not feed on molluscs can cause limited jumping or valve-closing reactions. The swimming action is performed by rapidly clapping the valves and expelling jets of water from each side of the hinge so that it moves with the curved edge of the shell at the front. The scallop jumps forwards by gradually relaxing the adductor muscle and then rapidly opening and closing the valves. Pecten maximus tends to be more numerous in areas where they are not fully exposed to strong currents. Scallops which live in sheltered habitats grow faster than scallops in areas exposed to wave action, possibly due to the filter feeding apparatus being unable to function because of high concentrations of particulate matter in the water in areas subject to high levels of wave exposure. Another factor that may be significant is that the processes of larval settlement and byssal attachment are rather delicate and would be disturbed in strong currents. Abundance and growth rates are negatively correlated with the amount of mud in the substrate. Scallops use larval motility to distribute themselves, as the adult scallops have limited mobility. The distribution of the larvae is affected by factors such as local hydrographic regimes and their survival, and this results in the scallops having an aggregated distribution within their geographic range. This means that the major fishing grounds are normally widely separated and each fishing ground's environmental conditions mean there are marked differences in structures of the populations, although the genetics of scallops are rather uniform across its range.