Aquilegia

Aquilegia (common names: granny's bonnet, columbine) is a genus of about 60–70 species of perennial plants that are found in meadows, woodlands, and at higher altitudes throughout the Northern Hemisphere, known for the spurred petals of their flowers. The genus name Aquilegia is derived from the Latin word for eagle (aquila), because of the shape of the flower petals, which are said to resemble an eagle's claw. The common name 'columbine' comes from the Latin for 'dove', due to the resemblance of the inverted flower to five doves clustered together. The leaves of this plant are compound and the flowers contain five sepals, five petals and five pistils. The fruit is a folliclewhich holds many seeds and is formed at the end of the pistils. Underneath the flower are spurs which contain nectar, mainly consumed by long-beaked birds such as hummingbirds. Columbines are closely related to plants in the genera Actaea (baneberries) and Aconitum (wolfsbanes/monkshoods), which like Aquilegia produce cardiogenic toxins. They are used as food plants by some Lepidoptera (butterfly and moth) caterpillars. These are mainly of noctuid moths – noted for feeding on many poisonous plants without harm – such as cabbage moth (Mamestra brassicae), dot moth (Melanchra persicariae) and mouse moth (Amphipyra tragopoginis). the engrailed (Ectropis crepuscularia), a geometer moth, also uses columbine as a larval food plant. The larvae of the Papaipema leucostigma also feed on columbine. Plants in the genus Aquilegia are a major food source for Bombus hortorum, a species of bumblebee. Specifically, they have been found to forage on species of Aquilegia vulgaris in Belgium and Aquilegia chrysantha in North America and Belgium. The bees do not show any preference in color of the flowers. Columbine is a hardy perennial, which propagates by seed. It will grow to a height of 15 to 20 inches. It will grow in full sun; however, it prefers growing in partial shade and well drained soil, and is able to tolerate average soils and dry soil conditions. Columbine is rated at hardiness zone 3 in the United States so does not require mulching or protection in the winter. Large numbers of hybrids are available for the garden, since the European A. vulgaris was hybridized with other European and North American varieties. Aquilegia species are very interfertile, and will self-sow. Some varieties are short-lived so are better treated as biennials. The following hybrid cultivars have gained the Royal Horticultural Society's Award of Garden Merit: