The black swan (Cygnus atratus) is a large waterbird, a species of swan which breeds mainly in the southeast and southwest regions of Australia. Within Australia they are nomadic, with erratic migration patterns dependent upon climatic conditions. Black swans are large birds with mostly black plumage and red bills. They are monogamous breeders, and are unusual in that one-quarter of all pairings are homosexual, mostly between males. Both partners share incubation and cygnet rearing duties. Black swans were introduced to various countries as an ornamental bird in the 1800s, but have escaped and formed stable populations. A small population of black swans exists on the River Thames at Marlow, on the Brook running through the small town of Dawlish in Devon (they have become the symbol of the town), near the River Itchen, Hampshire, and the River Tees near Stockton on Tees. Described scientifically by English naturalist John Latham in 1790, the black swan was formerly placed into a monotypic genus, Chenopis. Black swans can be found singly, or in loose companies numbering into the hundreds or even thousands. Black swans are popular birds in zoological gardens and bird collections, and escapees are sometimes seen outside their natural range. Black swans are mostly black-feathered birds, with white flight feathers. The bill is bright red, with a pale bar and tip; and legs and feet are greyish-black. Cobs (males) are slightly larger than pens (females), with a longer and straighter bill. Cygnets (immature birds) are a greyish-brown with pale-edged feathers. A mature black swan measures between 110 and 142 centimetres (43 and 56 in) in length and weighs 3.7–9 kilograms (8.2–19.8 lb). Its wing span is between 1.6 and 2 metres (5.2 and 6.6 ft). The neck is long (relatively the longest neck among the swans) and curved in an 'S'-shape. The black swan utters a musical and far reaching bugle-like sound, called either on the water or in flight, as well as a range of softer crooning notes. It can also whistle, especially when disturbed while breeding and nesting. When swimming, black swans hold their necks arched or erect and often carry their feathers or wings raised in an aggressive display. In flight, a wedge of black swans will form as a line or a V, with the individual birds flying strongly with undulating long necks, making whistling sounds with their wings and baying, bugling or trumpeting calls. The black swan is unlike any other Australian bird, although in poor light and at long range it may be confused with a magpie goose in flight. However, the black swan can be distinguished by its much longer neck and slower wing beat. One captive population of black swans in Lakeland, Florida has produced a few individuals which are a light mottled grey colour instead of black.