Abstract Sheep of a line (S) selected on an index to increase lean weight and decrease fatness at an age, and a control line (C), were given a high quality food at different levels including ad libitum. Live performance was measured from about 21 to 114 kg live weight. The carcasses of each line were analysed for lean, fat and bone at three widely varying weights in both males and females. Level of feeding did not affect the extent to which S was superior to C in either the level of fatness in the carcass (0·86 as much) or the ratio of lean to fat (1·28 as much). The lean to bone ratio was slightly greater in S (1·028 of the value of C; P 0·05) and was higher on the lowest level of feeding compared with the two higher levels used (P 0·05 in one experiment on females and P 0·001 in another on males). On ad libitum feeding the S line grew 1·19 times as fast and was 1·17 times as efficient compared with C. These advantages to S decreased as level of feeding decreased to become virtually zero at the lowest level of feeding used, which allowed C to grow at only 0·53 of the rate seen on ad libitum feeding. On ad libitum feeding growth was well described by a Gompertz growth function of the form W = (Z/B) exp(-exp (G0 –B t)). The maximum growth rate is (Z/e). Line S had a value of Z that was 1·10 that of C averaged across the two sexes. A Spillman function W = W0 + (A-W0) (1-exp (-k F)) was used to describe weight, W, in terms of cumulative intake, F. It worked well for ad libitum feeding and for the two restricted regimes used. The value of the combined parameter (A k) varied across treatments in the same way as efficiency did.
Abstract Two pig breeds, one improved (Cotswold Fl hybrid Large White × Landrace pigs = LWX) and the other unimproved (Chinese Meishan pigs = CM) were used to test the proposition that the genotype of the pig has an effect on the selection of a diet from two foods that differ in their crude protein content. From 21 to 34 kg live weight, the pigs were given access to either one of three foods or a choice of two foods with similar digestible energy concentration (16 MJ digestible energy per kg) but a different crude protein (CP) concentration. This resulted in four dietary treatments: (i) free and continuous access to low (L) crude protein food alone (130 g CP per kg, no. = 4 of each breed); (ii) free and continous access to high (H) crude protein food alone (252 g CP per kg, no. - 4 of each breed); (Hi) free and continuous access to moderate (M) crude protein food alone (206 g CP per kg, no. = 4 of each breed) and (iv) free and continuous access to both foods L and H as a choice (no. = 6 of each breed). On all treatments the LWX performed significantly better than the CM pigs in terms of live-weight gain and food conversion efficiency ( P < 0·001). The LWX and CM pigs given access to a single food contained the same amounts of protein in their bodies at 34 kg live weight, but the CM pigs had a considerably higher lipid ( P < 0·001) and a lower water content ( P < 0·001). When given a choice, the LWX pigs selected a significantly higher proportion of foodH(521 v. 226 (s.e.d. 49) g food H per kg for LWX and CM respectively) and therefore, a higher CP content in their diet (194 v. 144 (s.e.d. 5·4) g CP per kg respectively) than the CM pigs. The performance of pigs given a choice between two foods, in terms of live weight and rate of protein gain, was comparable with the best performance achieved on a single food (M) for the LWX, and better than the best performance on a single food (L) for the CM pigs. Thus, when given a choice between an appropriate pair of foods that differ in their crude protein content, pigs are able to select a diet that meets their requirements and allows them to express the growth characteristics typical for their breed (genotype).
Abstract The hypotheses tested were that the expected preference of sheep for a food with adequate rumen degradable protein (RDP) supplemented with urea would be reduced both by the addition of a buffer (sodium bicarbonate (SB)) and by offering ad libitum access to hay. A control food (C), calculated to be adequate in its ratio of effective RDP to fermentable metabolizable energy (fME), was formulated. Other foods were made by adding 12·5 (U 1 ) or 25 (U 2 ) g urea per kg fresh matter (FM) to C and 20 g SB per kg FM to C, U 1 and U 2 . The acid buffering capacity (ABC) of each food was measured in vitro. The experiment consisted of two successive periods, each of 4 weeks. Ninety-eight female, Texel ✕ Greyface sheep were randomly allocated to 14 groups each with seven animals. Groups 1 to 6 were offered one of: C, U 2 , C + SB, U 2 + SB, C with hay or U 2 with hay throughout the experiment. Groups 7 to 10 were offered the choices of C v. U 1 or C v. U 2 , either with or without hay in a change-over design; animals that received hay during period 1 (groups 8 and 10) did not do so during period 2 and vice versa (groups 7 and 9). Groups 11 to 14 (no. = 7) were offered the choices of C v. U 1 or C v. U 2 , either with or without SB supplemented to both foods, in a change-over design. Adding either urea, or SB, or both to C had no effects on intake or live-weight gain when offered alone. Both supplements significantly (P 0·001) increased the ABC of food C. Throughout the experiment hay consumption was very low (overall mean: 23 (s.e. 2·5) g hay per sheep day). Offering hay caused no change in the preference for the urea-supplemented foods. Sheep offered the choices C v. U 1 or C v. U 2 , with neither hay nor SB, selected 0.466 (s.e. 0·036) and 0.588 (s.e. 0·025) kg/kg total food intake (TFI) of U 1 and U 2 respectively. The proportions of the urea-supplemented foods were significantly reduced (P 0.01) by SB supplementation: to 0.348 (s.e.0·045) and 0·406 (s.e.0·059) kg/kg TFI of U 1 and U 2 respectively. The effect of SB addition on the diet selection of sheep could be due to its buffering properties. When SB is added to both foods the need for urea to be used as a buffer is reduced with a consequent decrease in the proportion selected as the urea-supplemented food. Effects of diet on buffering may override other diet selection objectives, such as the avoidance of an excess intake of RDP.
Ten Large White × Landrace boars at an initial live weight of 43 kg were given free access to two isoenergetic foods with 119 (L) and 222 (H) g crude protein (CP) per kg food as a choice, for 54 days. Six more similar boars were given access to food H only. The diet selected by the choice-fed pigs (measured as the proportion of food H per kg total food intake) changed systematically with time; the CP content of the selected diet fell from 193 in the first 7 days to 146 g CP per kg food in the last 7 days. The performance of the choicefed pigs, over the 54-day period, was as good as that of those given food H only: live-weight gain was 1101 v. 1069 g/day and food conversion efficiency 0·380 v. 0·374 g gain per g food respectively. However, the choice-feeding system allowed boars to reduce the protein content of their diet as they grew. It is suggested that it might be possible to give pigs a choice between two appropriate foods during the whole growing-fattening period instead of frequently changing the composition for their single food in order to try to meet their requirements.
It is frequently assumed that energy intakes from mixed foods with a high proportion of silage (HS) are lower than those from mixed foods with a high proportion of concentrate (HC), because of short-term constraints, i.e. gut fill, that physically limit the amount of food a cow can consume. It was the aim of the present study to analyse how different proportions of concentrate in mixed foods affect short-term feeding behaviour. We hypothesised that cows offered HS are likely to have more meals that are more spread out during the day and vary less in size than cows offered HC. Alternatively, we expected higher correlations between meal size and the length of intervals before (pre-prandial) or after (post-prandial) meals for cows offered HS than for cows offered HC. We tested the hypotheses with a data set of 21195 meals.
Abstract 1. An experiment was conducted to measure the potential growth of males and females of 6 commercial broiler stocks, from which information the growth rates of these genotypes could be characterised by the Gompertz growth equation. 2. Feeding and environmental conditions were designed to ensure that the birds remained comfortable throughout their growing period, which was to 26 weeks of age. A choice of diets differing in protein content was offered from 3 weeks of age. Because of leg weaknesses among the male broilers after 11 weeks of age, and because many females reached sexual maturity at about this age, the growth analyses were conducted on weights collected up to 11 weeks of age only. At this weight, broilers had achieved approximately 0–76 of their mature weight. 3. Birds representative of each genotype were killed for carcase analysis at weekly intervals to 9 weeks of age, and every two weeks thereafter. The contents of gut fill, feathers, water, protein, ash and lipid were measured on each of these birds; from these, equations were derived for each genotype that allowed the estimation of the weights of these components in the birds remaining on the experiment. 4. The body weight, body protein, body water and feather weight of the 12 genotypes were described in terms of the mature weight of these components, their rates of maturing and the time taken to reach the maximum rate of growth of each component. These descriptors of the growth of each component were then compared between genotypes. 5. No statistically significant differences existed in the rates of maturing of the different genotypes, either between strains or between sexes. Highly significant differences were evident between strains and between sexes in their mature weights, indicating that their rates of growth differed. 6. Estimates of mature feather weights indicated that this component of the body comprised 0.062 and 0.050 of the mature body weight of female and male broilers respectively. The protein content of feathers increased steadily, and the water content decreased steadily, throughout the growing period. 7. Differences between the genotypes evaluated in this experiment indicate that the nutrient and environmental requirements of these genotypes would differ. A description of each genotype, therefore, is an essential component of any simulation model that attempts to determine the optimum economic feeding programme and environmental conditions for broilers. Notes To whom correspondence should be addressed.
Selection in commercial populations on aspects of output, such as for growth rate in poultry, against fatness and for growth rate in pigs, and for milk yield in cows, has had very large effects on such outputs over the past 50 years. Partly because of the cost of recording intake, there has been little or no selection for food intake or feeding behaviour. In order to predict the effects of such past, and future, selection on intake it is necessary to have some suitable theoretical framework. Intake needs to be predicted in order to make rational feeding and environmental decisions. The idea that an animal will eat ‘to meet its requirements’ has proved useful and continues to be fruitful. An important part of the idea is that the animal (genotype) can be described in a way that is sufficient for the accurate prediction of its outputs over time. Such descriptions can be combined with a set of nutritional constants to calculate requirements. There appears to have been no change in the nutritional constants under selection for output. Under such selection it is simplest to assume that changes in intake follow from the changes in output rates, so that intake changes become entirely predictable. It is suggested that other ways that have been proposed for predicting intake cannot be successful in predicting the effects of selection. Feeding behaviour is seen as being the means that the animal uses to attain its intake rather than being the means by which that intake can be predicted. Thus, the organisation of feeding behaviour can be used to predict neither intake nor the effects of selection on it.
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